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- Gene S controls the sharpness of spines in a type of cactus. Cactuses with the dominant allele, S, have sharp spines, whereas homozygous recessive ss cactuses have dull spines. At the same time, a second gene, N, determines whether cactuses have spines. Homozygous recessive nn cactuses have no spines at all17. A cross between a true-breeding sharp-spined cactus and a spineless cactus would produce a.all sharp-spined progenyb.50% sharp-spined, 50% dull-spined progenyc.25% sharp-spined, 50% dull-spined, 25% spineless progenyd.all spineless progenye.It is impossible to determine the phenotypes of the progeny