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C'est une succession de réaction d'oxydo-réduction, qui ont lieu au sein d'un complexe protéique, qui sont transmembranaire, que se soit un transfert spontané d'un réducteur puissant à un fort oxydant comme dans la chaine respiratoire mitochondriale ou un transfert non spontanée nécessitant la lumière comme la chaine photosynthétique.

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